• Taxonomy, systematics, phylogeny, evolution, development, and biogeography of marine bristle worms;

  • Ecology of soft-bottom benthic communities


PhD, 2008            P.P. Shirshov Institute of Oceanology, Russian Academy of Sciences

Thesis title: “Taxonomy, phylogeny and biogeography of Onuphidae (Annelida: Polychaeta)”.

Supervisor: Dr. A.V. Gebruk, scientific consultant: Dr. K. Fauchald

MSс, 2004             Moscow State University, Department of Invertebrate Zoology

Thesis title: “Deep-sea benthic macrofaunal communities of the northern part of the Greenland Sea”.

Supervisor: Prof. A.B. Tzetlin, co-supervisors: Dr. A.V. Gebruk, Dr. V.O. Mokievsky


2013–2017 Marie Curie Fellowship. “Progenetic evolution in the deep-sea fauna (PRODEEP)”. University Museum of Bergen, Norway; in collaboration with Dr. E. Willassen.


2013–2017 Research Council of Norway (top-up finding of Marie Curie fellows)


2012–2013 YGGDRASIL – Young Guest and Doctoral Researchers' Annual Scholarships for Investigation and Learning in Norway. Research Council of Norway. “Evolution of the bristle worm family Onuphidae (Polychaeta, Annelida)”  University Museum of Bergen, Norway; in collaboration with Dr. E. Willassen.


2012 Zoologica Scripta Travel Fellowship, co-funding to the YGGDRASIL grant.


2012 – 2013 Russian Foundation for Basic Research (RFBR) – 12-05-33049, “Colonisation of ocean depth and adaptations of deep-sea fauna". P.P. Shirshov Institute of Oceanology RAS, Moscow, Russia.


2012 Endeavour Research Fellowship."Diversity and taxonomy of the deep-sea Australian onuphid seaworms". Australian Museum, Sydney, Australia; in collaboration with Dr. P. Hutchings and Dr. H. Paxton.


2010–2012  Russian Foundation for Basic Research (RFBR) – 10-04-01687 “Taxonomy and evolution of polychaetes of the family Onuphidae Kinberg, 1865 and their placement within the classis Polychaeta (Annelida)”. P.P. Shirshov Institute of Oceanology RAS, Moscow, Russia.


2011 Lerner-Gray Fellowship (The American Museum of Natural History). "Reproduction and development of a tube dwelling polychaete Mooreonuphis stigmatis (Treadvell, 1922) (Onuphidae) from the North-east Pacific". Field project at the Friday Harbor Labs, University of Washington, USA; in colaboration with Dr. E. Vortsepneva.


2010 Census of the Diversity of Abyssal Marine Life (CeDAMar) taxonomic exchange fellowship. “Quill worms (Onuphidae, Polychaeta) from the deep-sea German expeditions to Antarctic and South Atlantic (ANDEEP I, II, III, DIVA 1, 2, 3)”. The German Centre of the Marine Biodiversity Research, Hamburg, Germany; in collaboration with Dr. Karin Meißner.


2006–2007 Predoctoral fellowship at the National Museum of Natural History, Smithsonian Institution “Revision of the genera Paradiopatra, Diopatra and Epidiopatra (Polychaeta: Onuphidae); in collaboration with Dr. Kristian Fauchald.


Polina Borisova M.Sc. student (2016-2018). B.Sc. theis  (2016). Moscow State University, Russia (co-supervisor Dr. A. Tzetlin)


Martin M. Hektoen M.Sc. thesis (2015). 

University of Bergen, Norway (co-supervisor Dr. E. Willassen)




2016     12th International Polychaete Conference, Cardiff, UK

2016     ForBio: Research School in Biosystematics annual

              meeting, Trondheim, Norway

2014     ForBio: Research School in Biosystematics annual

              meeting, Tromsø, Norway

2013     11th International Polychaete Conference, Sydney,


2013     International polychaete workshop, Lizard Island

              Research Station, Australia

2010     10th International Polychaete Conference, Lecce, Italy

2010     12th International Deep-Sea Biology Symposium,

              Reykjavik, Iceland

2007     9th International Polychaete Conference, Portland,

              Maine, USA

2007     Abyssal Polychaete International Workshop, London, UK

2006     11th International Deep-Sea Biology Symposium,

              Southampton, UK



2013-      ForBio: Research School in Biosystematics

2011-      The Systematics Association

2011-      The Willi Hennig Society


2013        Collecting trip, Lizard Island Research Station                                      (Australian Museum, Sydney)

2013        Collecting trip, Mallorca, Spain

2012        Cruise on the RV Polarstern ARK XXVII/2, Greenland

                 Sea, HAUSGARTEN (AWI, Germany)

2009        Cruise on the RRS James Cook, Mid-Atlantic Ridge

                 (ECOMAR, NOCS, UK)

2007        Collecting trip, Espeland Marine Biological Station

                 (University of Bergen, Norway)

2006        Collecting trip at the Carrie Bow Cay Field Station

                 (Smithsonian Institution, Belize)

2005        Cruise on the RV Akademik Mstislav Keldysh (IO RAS),

                 Mid-Atlantic Ridge hydrothermal vents

2004        Cruise on the RV Rift (IO RAS), Caspian Sea

2003        Cruise on the RV Polarstern ARK XIX/3c to the Greenland

                 Sea, HAUSGARTEN (AWI, Germany)

2000 –     Yearly 1-2 month sampling trips at the White Sea

2011        Biological Station (MSU)


2013 – 2017           Marie Curie postdoctoral researcher at the Natural History Collections, University Museum of Bergen

2012 – 2013           Guest researcher at the Natural History Collections, University Museum of Bergen

2005 – 2013           Researcher at the Laboratory of Ocean Benthic Fauna, P.P. Shirshov Institute of Oceanology, RAS

2006 - 2007            Predoctoral fellow at the Natural History Musuem, Smithsonian Institution

2004 – 2005           Technician at the Laboratory of Ocean Benthic Fauna, P.P. Shirshov Institute of Oceanology, RAS